King George V (KG5) began convoy escort duties in February 1941. The ship also took part in an unsuccessful search for the German warships Scharnhorst and Gneisenau during the Kriegsmarine Operation Berlin.

The ship was made the flagship of the Home Fleet under the command of Admiral Sir John Tovey after the destruction of HMS Hood, and was involved in the chase and the sinking of Hood's opponent, the German battleship Bismarck. On 27 May 1941, she fired 339 x 14 inch and 660 x 5.25 inch shells at Bismarck. King George's shell fire helped damage Bismarck's Superstructure and disable its main armament. At 10:39 that morning, Bismarck sank.

While escorting convoy PQ-15 to Murmansk on 1 May 1942, King George V collided with the destroyer HMS Punjabi, resulting in the sinking of the latter ship with 49 crew, and bow damage to the battleship.

In the Mediterranean, King George V covered the Operation Husky landings at Sicily, as well as transporting the Prime Minister, Winston Churchill, back to Britain from the Tehran Conference.

From 1944 until the surrender of Japan, King George V served with the British Pacific Fleet, and was present off Japan during the official surrender ceremony.

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